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Alex Greiner, M5G


MSTP Learning Community: Boulware

Advanced Standing Entry:  June 15, 2016

Matriculation to CCOM:  2014

PhD Program:



Barry London, MD, PhD

Genetic and Molecular Basis of Sudden Cardiac Death Syndromes

Thesis Defense:  January 28, 2022


2014 - MOHDI liason committee

2015-2018 - MSTP Wellness Committee

2016-2017 - MSTP Applicant Task Force

2017-2020 - MSTP Admissions Committee Member

2019 - MSTP RAGBRAI Planning Team


2015 - Medical Student Research Day-Research Excellence in Internal Medicine

2017 - Microfunding ($5k) grant from the Pappajohn Biomedical Institute to study the contributions of mutations in DEPDC5 to sudden cardiac death

2018-2018 - Pre-doctoral fellowship from the American Heart Association

2018-2022 - NIH F30 Fellowship, National Heart, Lung & Blood Institute

Individual Funding Awards:

09/17/2018 - 09/16/2022
F30 HL143908 - (National Heart, Lung & Blood Institute)
Assessing the Impact of MSX1 in Cardiac Fibrosis and Sudden Cardiac Death

07/01/2018 - 09/17/2018
American Heart Association
Assessing the Role of MSX1 in Cardiac Fibrosis and Sudden Cardiac Death (Declined due to F30 funding)


2019 Fall - Pathogenesis of Metabolic and Cardiovascular Disease, Program: Molecular Medicine


2014 - BS, Chemistry - Luther College


ORCiD:  0000-0003-3082-2091

London B, Greiner AM, Mehdi H, Gutmann R. IDENTIFYING NEW SUDDEN DEATH GENES. Trans Am Clin Climatol Assoc. 2018;129:183-184. PubMed PMID: 30166713; PubMed Central PMCID: PMC6116611.

Greiner AM, Kaldjian LC. Rethinking medical oaths using the Physician Charter and ethical virtues. Med Educ. 2018 Aug;52(8):826-837. doi: 10.1111/medu.13581. Epub 2018 Apr 27. PubMed PMID: 29700846.

London B, Greiner AM, Mehdi H, Gutmann R. Identifying New Sudden Death Genes. Trans Am Clin Climatol Assoc. 2018;129:183-184. PMID: 30166713; PMCID: PMC6116611.

Matasic DS, Yoon JY, McLendon JM, Mehdi H, Schmidt MS, Greiner AM, Quinones P, Morgan GM, Boudreau RL, Irani K, Brenner C, London B. Modulation of the cardiac sodium channel NaV1.5 peak and late currents by NAD+ precursors. J Mol Cell Cardiol. 2020 Apr;141:70-81. doi: 10.1016/j.yjmcc.2020.01.013. Epub 2020 Mar 21. PubMed PMID: 32209328; PubMed Central PMCID: PMC7234910.

Greiner A, Mehdi H, Cevan C, Gutman R, London B.  Additional Evidence Implicating GPDIL in the Pathogenesis of Brugada Syndrome in a Large Multi-generational Family.  medRxiv 2022.09.17.22280058; https://doi.org/10.1101/2022.09.17.22280058.