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The Department of Pathology Master of Science (MS) Graduate Program prepares post-baccalaureate science majors for a range of biomedical careers.
Graduate-level coursework provides graduate students with a foundation in cellular and molecular biology, as well as specialized knowledge in pathobiology. A laboratory intensive thesis project equips trainees with cutting edge research skills. The Pathology MS Program is designed for graduates to advance into research assistant or research scientist positions in academic and private sector laboratories, or be competitive for advanced degree (PhD, MBA or MD) programs.

Admissions
Application Process
The application process is paperless, with all application materials either entered or uploaded onto the admissions portal. The web portal for application can be accessed at the Graduate Admissions Application Portal.
In addition to providing basic information, applicants will be asked to upload:
- Transcripts from all undergraduate or graduate institutions attended.
- A statement of purpose that summarizes your academic career to date, all research experiences, a brief narrative as to why you wish to pursue an M.S. in Pathology, and your future goals.
- A current CV or résumé. It should chronologically list your educational, research and work experience to date. The CV/résumé should also list any awards, conferences, presentations and publications. Applicants are welcome to include items such as volunteering, study abroad experiences, teaching assignments and other achievements.
- Three letters of recommendation are required, and it is helpful if those writing letters are able to comment on your laboratory experience and/or aptitude for basic research. You will be asked to provide the contact information (email address) for three references, and they will be instructed to upload their letters. Letters must be submitted on official letterhead and signed/authenticated by the letter writer.
For more information, please read: Pathology MS Program Admission Requirements
Qualifications
Applicants must have a bachelor’s degree in a science discipline from a Regionally Accredited American College or University, or an equivalent degree from another country as determined by the Office of Admissions. Applicants must also have a minimum grade-point average (GPA) of 3.00/4.00, or the foreign equivalent as determined by the Office of Admissions. In addition, laboratory-based research experience is highly desired. Finally, international students must submit English proficiency test scores that meet institutional requirements. For more information on English proficiency requirements, please go read: Graduate College English Proficiency Requirements.
A subset of applicants will be selected for interviews, conducted either in person or by Zoom. Admission decisions are determined based on academic performance, research experience, letters of recommendation, personal statement and interviews.
Deadlines and fees
Fall semester admission: January 15 for international applicants, February 15 for U.S. applicants.
Spring semester admission: September 15 for international applicants; October 15 for U.S. applicants. Please note: Openings for spring semester admission are rare but may exist from time to time. Prior to applying, please inquire as to whether there are any spring openings.
The application fee for domestic applicants is $60 and $100 for international applicants. Applicants may be eligible for a Graduate Application Fee Waiver based upon undergraduate/post-baccalaureate program participation or financial hardship.

Program overview
Coursework
All Pathology M.S. students must have a minimum of 21 credit hours of coursework and 9 hours of research. Required coursework is as follows:
- BMED:5207:000A Principles Molecular & Cellular Biology (3 s.h.)
- BIOS:4120:0AAA Introduction to Biostatistics (3 s.h.)
- PATH:6220:0001 Seminar in Pathology (1 s.h. x 2 semesters)
- PATH:5270:0001 Pathogenesis of Major Human Diseases (3 s.h.)
- BMED:7270:0002 Principles of Scholarly Integrity
Required coursework results in:
- A basic understanding of molecular and cellular biology.
- A basic understanding of biostatistics.
- An advanced understanding of pathobiology and mechanisms of human disease.
Remaining coursework consists of electives focused on the area or topic of the student's thesis project. These electives are graduate level courses offered by a range of departments on the biomedical campus. All coursework is completed within the first two years.
Thesis project
The thesis project is carried out under the guidance of the mentor and thesis committee. The committee is composed of the student's mentor and two additional faculty members. In general, the thesis consists of four chapters with the first being a concise review of the literature, the second materials and methods, and the last two a scholarly description of the project results. The student must meet with the thesis committee at least once a year, with a satisfactory report submitted after each meeting. The thesis must be defended before the committee prior to final approval.
Faculty and laboratories
Who are our students?
Students admitted to the Pathology program consist of recent college graduates, as well as individuals who have been in the work force for several years.
Given the requirements for admission, the academic credentials of admitted Pathology students are on par with most biomedical PhD programs on campus. This often leads to questions as to why a student would apply to the Pathology MS rather than a PhD granting program.
Although our students are talented and science focused, most enter the program since they have not yet crystallized their long-term goals. As discussed below in Outcomes, the program is very successful in helping students realize their potential, plan careers and understand whether they wish to pursue an advanced degree.
Time to completion
Since 2005, the average time to completion has been 30 months (range = 21-39 months). Since 2010, average time to completion has dropped to 29 months (range = 21-39 months). Time to completion is measured by date of matriculation to date of thesis defense.
Outcomes
Since 2005, graduates of the program have used the M.S. degree to further their professional careers in a number of venues. Some have gone directly into the work force as research assistants or staff scientists in either academic or private sector laboratories. Other graduates have furthered their education by pursuing a range of advanced professional degrees. These include M.D., P.A., Ph.D., and executive M.B.A. programs. Since 2005, 74% of Pathology Program graduates have been accepted into advanced degree programs. To view the thesis projects and academic/employment history of Pathology Program graduates since 2005, please click on the link below:
Student resources
Stipend and tuition support
All Pathology graduate students receive full stipend and tuition support until they complete the program. Stipend amounts are at the same level as other graduate programs on the biomedical campus. Stipend support for the upcoming 12 month fiscal year (ending June 30, 2025) is $34,500. The stipend is paid to students on a monthly basis. Support for new incoming students begins on the first day of classes for the semester.
All Pathology graduate students have their tuition and fees paid until they complete the program.
The program also pays most of the costs for health and dental insurance. For more information on health and dental benefits, please read: Graduate Student Health and Dental Insurance.
Policy handbook
This policy handbook outlines all key information for students and faculty of the Department of Pathology Graduate Program:
Additional links and resources
Contact information
Program director

Vladimir Badovinac, PhD
Program oversight committee

Alexander Boyden, PhD

Kevin L. Legge, PhD

Ashutosh Mangalam, PhD

Munir Tanas, MD
Program coordinator

Gabriella Young, MLS
Current students

Maddisen Dwelley, BS

Lauryn Hahn, BS

Payton Kahl

Aracely Miron-Ocampo, BS

Ryan Swartz, MS
Alumni
Master of Science Program Alumni 2005–present
Pathology M.S. graduate | Previous Institution | Mentor/ Year of Thesis Defense | Thesis Project | After Graduation |
---|---|---|---|---|
Robin Goforth Williams | Indiana University | Dr. Sanjeev Sethi, 2005 | Vascular Injury to the Kidney: Association with Inherited Thrombophilia | M.D. UI Carver College of Medicine (2011) Pediatrics resident, University Iowa Pediatric Heme/Onc Fellow University of Minnesota, Minneapolis, MN Assistant Professor, Department of Pediatrics University of Minnesota |
Michelle Edsen Moore | Drake University | Dr. Annette Schlueter, 2005 | The Effects of Chronic Ethanol Exposure on Dendropoiesis and Costimulatory Molecule Expression | MBA, Univ of Iowa (2011) Program Associate, University of Iowa Dental School Client Services Senior Manager Beckman-Coulter, Inc. |
Cynthia Gunsolly | Loras College | Dr. Jonathon Heusel, 2006 | Transcriptional Control of an Unusual Receptor of Human Natural Killer Cells - KIR2DL4 | Research Assistant II, Pathology, University of Iowa CLS, St. Luke’s Hospital, Cedar Rapids, IA |
Kristin Ness | University of Wisconsin Madison | Dr. Annette Schlueter, 2006 | Chronic Ethanol Consumption Alters Dendritic Cell Migration | Ph.D. Immunology (2011), University of Iowa Post-doctoral Fellow, University of Iowa |
Alison Esser | Iowa State University | Dr. Michael Henry, 2006 | Endothelin-1 Biosynthesis and Function in Prostate Cancer Metastasis | Ph.D. Physiology (2011), University of Iowa Post-doctoral Fellow, Washington University, St. Louis Advising Coordinator, Genetics and Cell Bio, Iowa State University, Ames, IA |
Aja Davis Rich | Grinnell College | Dr. Jonathon Heusel, 2007 | Determinants of Virus-specific Natural Killer Cell Activation: Characterization and Mutational Analysis of the Murine Cytomegalovirus Glycoprotein m157 | MBA, Executive MBA program (2009), University of Virginia, Charlottesville Marketing Division, Amgen Corporation, Thousand Oaks, CA Marketing Manager, Baxter International, Deerfield, IL Senior Director, New Product Planning, GI and Neuroscience, Takeda, Chicago |
Jodi McGill | Iowa State University | Dr. Kevin Legge, 2007 | The Role of Pulmonary Dendritic Cells in the Lungs During Influenza Infection | Ph.D. Immunology (2010), University of Iowa Post-doctoral Fellow, Iowa State University, Ames, IA Assistant Professor, Kansas State University, Manhattan, KS Associate Professor, Iowa State University, Ames, IA |
Prajwal Gurung | Winona State University | Dr. Robert Cook, 2007 | Antigen-specific CD8+ T Cell Defects in Chronic Ethanol Administered Mice | Ph.D. Immunology (2011), University of Iowa Post-Doctoral Fellow, St. Jude Hospital, Memphis, TN Assistant Professor Associate Professor, University of Iowa |
Guinevere Lykken Strand | Luther College | Dr. Aaron Bossler, 2008 | PTPN13 is a Potential Tumor Suppressor for Head and Neck Squamous Cell Carcinoma | Instructor, Department of Biology, Luther College, Decorah, IA |
Gregory Thomas | University of Iowa | Dr. Hasem, Habelhah, 2008 | TRAF2 Phosphorylation at Serine 55 Regulates Secondary NF-κB Activity | Ph.D. Molecular and Cell Biology (2012) University of Iowa Post-Doctoral Fellow, University of Iowa Assistant Director of Cancer Career Enhancement and Training, Holden Comprehensive Cancer Center, University of Iowa |
Huy Nguyen | University of Washington, Seattle | Dr. Steven Moore, 2008 | Dystroglycan in Cerebellar Development and Disease | Ph.D. Neuroscience (2013), University of Iowa Post-Doctoral Fellow, Stanford University, Stanford, CA Principal Scientist, Surrozen Biotechnology, San Francisco, CA |
Corey Parlet | Concordia College, St. Paul | Dr. Annette Schlueter, 2009 | Chronic Ethanol Feeding Impedes the Acquisition of a Promigratory Phenotype in Murine Cutaneous Dendritic Cells | Ph.D. Immunology (2014), University of Iowa Post-Doctoral Fellow, University of Iowa Assistant Research Scientist, University of Iowa |
Caitlin Ross | University of Maine | Dr. Jackie Bickenbach, 2010 | Oxygen Tension Regulates Keratinocyte Migration in Aged Skin | Cytogenetic Lab Specialist, Department of Pediatrics, University Iowa M.D. University of Queensland/Ochsner Medical School (2015) Emergency Medicine Practice Queensland Health, Australia |
Deepa Rai | Winona State University | Dr. Vladimir Badovinac, 2010 | Tracking Total Polyclonal CD8 T Cell Responses in Inbred and Outbred Hosts | Research Specialist, Pathology, University of Iowa |
Khanh Duong | University of Iowa | Dr. Stacey Klutts, 2010 | Amylases and Aspergillus fumigatus Cell Wall Synthesis: New Roles for Classical Enzymes | Ph.D. Molecular and Cell Biology (2014), University of Iowa Post-Doctoral Fellow, Harvard Medical School, Boston, MA Lead Scientist, Homology Medicines, Inc., Bedford, MA Lead, New Technologies, Takeda Pharmaceuticals, Cambridge, MA |
Matthew Martin | Creighton University | Dr. Vladimir Badovinac, 2010 | Naive and Memory CD8 T Cell Responses After Antigen Stimulation In Vivo | Ph.D. Immunology (2015), University of Iowa Post-Doctoral Fellow, University of Iowa Research Scientist University of Minnesota, Minneapolis, MN Laboratory Leadership Service Fellow, CDC, Madison WI |
Danial Samar | University of California, Davis | Dr. Stacey Klutts, 2012 | Identification and Characterization of TFT1, a Glycosyltransferase Necessary for Cell Wall β-1,3-1,4-glucan Synthesis in Aspergillus fumigatus | Manager of Global Inside Sales Integrated DNA Technologies, Coralville, IA Director of Commercial Operations, Teknova Biotechnology, Eugene, OR |
Lea Goldman | Creighton University | Dr. Thomas Waldschmidt, 2013 | Kinetics and Phenotype of the Draining Lymph Node and Pulmonary B Cell Response to an Influenza Virus Like Particle Vaccine | D.O. Alabama College of Osteopathic Medicine (2017), Dothan, AL Ob/Gyn residency, Henry Ford, Macomb Hospital, Clinton, MI Ob/Gyn practice, Dothan, AL |
Kaylia Duncan | Fisk University | Dr. Siegfried Janz, 2013 | Dynamics of Tumor Progression and Therapy Response in IL-6 and Myc Driven Plasma Cell Malignancy | Ph.D. Molecular Medicine (2019), University of Iowa Post-Doctoral Fellow, University of Iowa Post-Doctoral Researcher, Children’s Hospital of Philadelphia (CHOP) |
Sean Duong | Loyola University | Dr. Vladimir Badovinac, 2013 | The Impact of Poly-Microbial Sepsis on Pre-Existing Memory CD8 T Cell Responses | Laboratory Supervisor, BioLife Plasma Services, Chicago, IL |
Nicholas Borcherding | Iowa State University | Dr. Weizhou Zhang, 2014 | Non-Canonical Wnt Signaling in Breast Cancer Initiation and Progression | M.D. and Ph.D. UI Carver College of Medicine (2020), University of Iowa Physician/Scientist Resident Trainee, Department of Pathology, Washington University, St. Louis |
Adam Koch | University of Nebraska | Dr. Andrean Simons-Burnett, 2014 | The Role of MyD88-dependent Signaling in the Anti-tumor Efficacy of the EGFR Inhibitor Erlotinib in Head and Neck Cancer | M.D. Loma Linda Medical School, Loma Linda, CA (2018) Internal Medicine Resident, Mayo Clinic, Rochester, MN Internal Medicine practice, Mayo Clinic |
Emma Hornick | St. Olaf College | Dr. Kevin Legge, 2014 | T Cell Immunity to Respiratory Infections | Ph.D. Immunology (2018), University of Iowa Post-Doctoral Fellow, University of Iowa |
Sarah Hall | Central Michigan University | Dr. Munir Tanas, 2017 | Characterizing the Contribution of Hippo Pathway Dysregulation in Sarcomagenesis | Research Laboratory Specialist, Michigan Center for Translational Pathology, University of Michigan Medical Center, Ann Arbor, MI |
Rachel Dahl | Winona State University | Dr. Andrean Simons-Burnett, 2018 | Targeting Interleukin-6 Trans-signaling in Head and Neck Squamous Cell Carcinoma | M.D. Carver College of Medicine (2022), University of Iowa Surgery Resident, Hennepin County, Medical Center, Minneapolis, MN |
Sung Jo | University of Florida | Dr. Weizhou, Zhang, 2018 | Targeting MSH2-MSH6 Heterodimer in Treating Basal-like Breast Cancer | Research Associate II, Morphic Therapeutic, Boston, MA Associate Scientist, Intellia Therapeutics, Cambridge, MA |
Khaliunaa Bayanbold | Thomas Jefferson University | Dr. Ling-Zhi Liu, 2019 | miR-27a/b Suppression and Nrf2 Upregulation Promote Cr(VI) Transformed Carcinogenesis and Angiogenesis | Ph.D. Free Radical and Radiation Biology Program, University of Iowa Post-Doctoral Fellow, University of Iowa |
Krista Thornton | Eastern New Mexico University | Dr. Siefried Janz, 2019 | The Role of Transcription Factor FOXM1 in High-Risk Multiple Myeloma | Human Resources Consultant Community Colleges of Spokane Spokane, WA |
Kimberly Fiock | University of Texas - Dallas | Dr. Marco Hefti, 2020 | The Developmental Expression and Regulation of Tau | Ph.D. Exp Pathology Program (2023), University of Iowa Research Scientist, University of Iowa |
Monisha MIttal | University of Delhi, India | Dr. Jian Zhang 2021 | Cyclic GMP-AMP Synthase Regulates the Development of Experimental Autoimmune Encephalomyelitis | M.S. Bioinformatics Program, Northeastern University, Boston, MA Computational Biologist II, UT Southwestern Medical Center, Dallas, TX |
Steven Moioffer | University of Iowa | Dr. Vladimir Badovinac, 2021 | Impact of Sepsis on Pathogen-specific Circulating and Resident Memory CD8 T Cell Subsets | Ph.D. student, Immunobiology Graduate Program, Yale School of Medicine, New Haven, CT |
Mohammad Heidarian | California State University, East Bay | Dr. Vladimir Badovinac, 2023 | Sublethal Whole-body Irradiation Induces Permanent Loss and Dysfunction in Pathogen-specific Circulating Memory CD8 T Cell Populations | Ph.D. student, Experimental Pathology Program, University of Iowa |
Peter Lehman | Colorado College | Dr. Ashutosh Mangalam, 2023 | Effect of Gut Microbiota Modulation During Early Life and Adulthood on Host Immunity | Ph.D. student, Experimental Pathology Program, University of Iowa |
Connor Wilhelm | University of Wisconsin, Madison | Dr. Nitin Karandikar, 2023 | Proteolipid Protein-induced Mouse Model of Multiple Sclerosis Requires B Cell-mediated Antigen Prestation | Ph.D. student, Experimental Pathology Program, University of Iowa Research Assistant, Department of Pathology, University of Iowa |
Romina Abbasian | California State University, East Bay | Dr. Dustin Bosch, 2023 | Defense Against Bacterial Toxin Secretion Systems | Research Associate, University of California, San Francisco |
Elizabeth Escue | Southern Illinois University Carbondale | Dr. Vladimir Badovinac, 2025 | The ability of memory CD8 T cell subsets to numerically and functionally recover following whole body irradiation is influenced by their history of cognate antigen exposures. |